How to Make It
Preheat the oven to 350°. Cook the penne in a large pot of boiling salted water, stirring occasionally, until al dente. Drain the pasta, reserving 1/2 cup of the cooking water. Return the pasta to the pot.
In a small bowl, toss the bread crumbs with the Parmesan and 2 tablespoons of the olive oil. In a large skillet, heat the remaining 1/4 cup of olive oil, add the garlic and cook over moderately low heat until golden, about 4 minutes. Stir in the tuna and scallions and remove from the heat. Add the olives, capers, lemon juice, oregano and crushed red pepper. Season with salt and black pepper.
Toss the penne with the tuna sauce, adding a few tablespoons of the pasta cooking water to moisten the pasta. Transfer to a 9-by-13-inch baking dish and sprinkle with the bread crumbs. Bake for 8 minutes, or until heated through. Sprinkle with the parsley and serve.